Selected publications

Wang, F., Ding, P., Liang, X., Ding, X., Brandt, C. B., Sjöstedt, E., Zhu, J., Bolund, S., Zhang, L., de Rooij, L., Luo, L., Wei, Y., Zhao, W., Lv, Z., Haskó, J., Li, R., Qin, Q., Jia, Y., Wu, W., Yuan, Y., … Luo, Y. (2022). Endothelial cell heterogeneity and microglia regulons revealed by a pig cell landscape at single-cell level. Nature communications, 13 (1), 3620. doi.org/10.1038/s41467-022-31388-z

Noer JB, Hørsdal OK, Xiang X, Luo Y, Regenberg B. Extrachromosomal circular DNA in cancer: history, current knowledge, and methods. Trends Genet. 2022 Mar 8:S0168-9525(22)00034-8. doi: 10.1016/j.tig.2022.02.007. Epub ahead of print. PMID: 35277298. (This review provides a comprehensive summary of the eccDNA and its roles in cancers.)

Corsi GI, Qu K, Alkan F, Pan X, Luo Y, Gorodkin J. CRISPR/Cas9 gRNA activity depends on free energy changes and on the target PAM context. Nat Commun. 2022 May 30;13(1):3006. doi: 10.1038/s41467-022-30515-0. PMID: 35637227; PMCID: PMC9151727.

Karlsson M, Sjöstedt E, Oksvold P, Sivertsson Å, Huang J, Álvez MB, Arif M, Li X, Lin L, Yu J, Ma T, Xu F, Han P, Jiang H, Mardinoglu A, Zhang C, von Feilitzen K, Xu X, Wang J, Yang H, Bolund L, Zhong W, Fagerberg L, Lindskog C, Pontén F, Mulder J, Luo Y, Uhlen M. Genome-wide annotation of protein-coding genes in pig. BMC Biol. 2022 Jan 25;20(1):25. doi: 10.1186/s12915-022-01229-y. PMID: 35073880; PMCID: PMC8788080. (The first body-wide classification of protein coding gene expression in pig tissues and organs)

Xiang X, Corsi GI, Anthon C, Qu K, Pan X, Liang X, Han P, Dong Z, Liu L, Zhong J, Ma T, Wang J, Zhang X, Jiang H, Xu F, Liu X, Xu X, Wang J, Yang H, Bolund L, Church GM, Lin L, Gorodkin J, Luo Y. Enhancing CRISPR-Cas9 gRNA efficiency prediction by data integration and deep learning. Nat Commun. 2021 May 28;12(1):3238. doi: 10.1038/s41467-021-23576-0. PMID: 34050182; PMCID: PMC8163799. (This study reports the CRISPRTRAP-seq method for high throughput quantification of CRISPR gRNA activity in cells. The study also reported the generation of new deep learning based tool, CRISPRon, for prediction of CRISPR on-target targeting activity.)

Karlsson M, Zhang C, Méar L, Zhong W, Digre A, Katona B, Sjöstedt E, Butler L, Odeberg J, Dusart P, Edfors F, Oksvold P, von Feilitzen K, Zwahlen M, Arif M, Altay O, Li X, Ozcan M, Mardinoglu A, Fagerberg L, Mulder J, Luo Y, Ponten F, Uhlén M, Lindskog C. A single-cell type transcriptomics map of human tissues. Sci Adv. 2021 Jul 28;7(31):eabh2169. doi: 10.1126/sciadv.abh2169. PMID: 34321199; PMCID: PMC8318366. (A single cell type gene expression atlas in human tissues. This study integrate single cell RNA sequencing results to classify the gene expression specificity in single cell type levels)

Sjöstedt E, Zhong W, Fagerberg L, Karlsson M, Mitsios N, Adori C, Oksvold P, Edfors F, Limiszewska A, Hikmet F, Huang J, Du Y, Lin L, Dong Z, Yang L, Liu X, Jiang H, Xu X, Wang J, Yang H, Bolund L, Mardinoglu A, Zhang C, von Feilitzen K, Lindskog C, Pontén F, Luo Y, Hökfelt T, Uhlén M, Mulder J. An atlas of the protein-coding genes in the human, pig, and mouse brain. Science. 2020 Mar 6;367(6482):eaay5947. doi: 10.1126/science.aay5947. PMID: 32139519. (This study reported the first systematic classification of gene expression specificity and distribution of all protein-coding genes in the mouse, pig and human brain.)

Kalucka J, de Rooij LPMH, Goveia J, Rohlenova K, Dumas SJ, Meta E, Conchinha NV, Taverna F, Teuwen LA, Veys K, García-Caballero M, Khan S, Geldhof V, Sokol L, Chen R, Treps L, Borri M, de Zeeuw P, Dubois C, Karakach TK, Falkenberg KD, Parys M, Yin X, Vinckier S, Du Y, Fenton RA, Schoonjans L, Dewerchin M, Eelen G, Thienpont B, Lin L, Bolund L, Li X, Luo Y, Carmeliet P. Single-Cell Transcriptome Atlas of Murine Endothelial Cells. Cell. 2020 Feb 20;180(4):764-779.e20. doi: 10.1016/j.cell.2020.01.015. Epub 2020 Feb 13. PMID: 32059779. (This study reported the first investigation of EC heterogeneity in mouse tissues/organs using single cell transcriptome analysis)

Niu D, Wei HJ, Lin L, George H, Wang T, Lee IH, Zhao HY, Wang Y, Kan Y, Shrock E, Lesha E, Wang G, Luo Y, Qing Y, Jiao D, Zhao H, Zhou X, Wang S, Wei H, Güell M, Church GM, Yang L. Inactivation of porcine endogenous retrovirus in pigs using CRISPR-Cas9. Science. 2017 Sep 22;357(6357):1303-1307. doi: 10.1126/science.aan4187. Epub 2017 Aug 10. PMID: 28798043; PMCID: PMC5813284. (The first PERV inactivated pigs were generated by CRISPR and cloning.)

Full publications

Zou, L., Luo, Y., Chen, M., Wang, G., Ding, M., Petersen, C. C., Kang, R., Dagnaes-Hansen, F., Zeng, Y., Lv, N., Ma, Q., Le, D. Q. S., Besenbacher, F., Bolund, L., Jensen, T. G., Kjems, J., Pu, W. T. & Bünger, C. (2013). A simple method for deriving functional MSCs and applied for osteogenesis in 3D scaffolds. Scientific Reports, 3, Article 2243. https://doi.org/10.1038/srep02243

Zhou, Y., Liu, Y., Hussmann, D., Brøgger, P., Al-Saaidi, R. A., Tan, S., Lin, L., Petersen, T. S., Zhou, G. Q., Bross, P., Aagaard, L., Klein, T., Rønn, S. G., Pedersen, H. D., Bolund, L., Nielsen, A. L., Sørensen, C. B. & Luo, Y. (2016). Enhanced genome editing in mammalian cells with a modified dual-fluorescent surrogate system. Cellular and Molecular Life Sciences, 73(13), 2543-2563. https://doi.org/10.1007/s00018-015-2128-3

Zhou, Y., Al-Saaidi, R. A., Guerra, P. F., Freude, K. K., Olsen, R. K. J., Jensen, U. B., Gregersen, N., Hyttel, P., Bolund, L., Aagaard, L., Bross, P. & Luo, Y. (2017). Mitochondrial Spare Respiratory Capacity Is Negatively Correlated With Nuclear Reprogramming Efficiency. Stem Cells and Development, 26(3), 166-176. https://doi.org/10.1089/scd.2016.0162

Zhou, X., Rahimi, K., Kjems, J. & Lin, L. (2023). High-resolution RNA isoform variation analysis by combining the advantages of single-cell RNA and nanopore sequencing. 44. Abstract from Neuroscience Day, Aarhus, Denmark.

Zhou, M., Lv, W., Han, P., Sun, K., Hao, Z., Gao, L., Xu, Y., Xu, Z., Shao, S., Ma, S., Guo, Q., Zhang, H., Liu, K., Yang, F., Yuan, Z., Wu, G., Yu, C., Luo, Y., Yao, Z. & Zhao, J. (2024). Plasma extrachromosomal circular DNA as a potential diagnostic biomarker for nodular thyroid disease. Clinical and Translational Medicine, 14(6), e1740. https://doi.org/10.1002/ctm2.1740

Zhou, T., Zhou, Y., Cömert, C., Zhou, X.-Y., Lin, L., Bolund, L., Palmfeldt, J., Luo, Y., Bross, P. & Tong, G. (2026). Energy metabolism alteration and gene expression reprogramming in a cell model of high fat load non-alcoholic fatty liver disease. BMC Gastroenterology, 26(1), Article 160. https://doi.org/10.1186/s12876-026-04658-z

Zheng, T., Huang, J., Xiang, X., Li, S., Yu, J., Qu, K., Xu, Z., Han, P., Dong, Z., Liu, Y., Xu, F., Yang, H., Jäättelä, M., Luo, Y. & Liu, B. (2021). Systematical analysis reveals a strong cancer relevance of CREB1-regulated genes. Cancer Cell International, 21(1), Article 530. https://doi.org/10.1186/s12935-021-02224-z

Zhao, X., Qu, K., Curci, B., Yang, H., Bolund, L., Lin, L. & Luo, Y. (2023). Comparison of In-Frame Deletion, Homology-Directed Repair, and Prime Editing-Based Correction of Duchenne Muscular Dystrophy Mutations. Biomolecules, 13(5), Article 870. https://doi.org/10.3390/biom13050870

Zhang, J., Li, H., Teng, H., Zhang, T., Luo, Y., Zhao, M., Li, Y.-Q. & Sun, Z. S. (2012). Regulation of Peripheral Clock to Oscillation of Substance P Contributes to Circadian Inflammatory Pain. Anesthesiology News, 117(1). https://doi.org/10.1097/ALN.0b013e31825b4fc1

Zhang, Y., Schmid, B., Nikolaisen, N. K., Rasmussen, M. A., Aldana, B. I., Agger, M., Calloe, K., Stummann, T. C., Larsen, H. M., Nielsen, T. T., Huang, J., Xu, F., Liu, X., Bolund, L., Meyer, M., Bak, L. K., Waagepetersen, H. S., Luo, Y., Nielsen, J. E. ... FReJA Consortium (2017). Patient iPSC-Derived Neurons for Disease Modeling of Frontotemporal Dementia with Mutation in CHMP2B. Stem Cell Reports, 8(3), 648-658. https://doi.org/10.1016/j.stemcr.2017.01.012

Zhang, C., Li, C., Yang, L., Leng, L., Jovic, D., Wang, J., Fang, F., Li, G., Zhao, D., Li, X., Lin, L., Luo, Y., Bolund, L., Huang, J., Lin, G. & Xu, F. (2021). The Dynamic Changes of Transcription Factors During the Development Processes of Human Biparental and Uniparental Embryos. Frontiers in Cell and Developmental Biology, 9, Article 709498. https://doi.org/10.3389/fcell.2021.709498

Zeng, Y., Wang, A., Lv, W., Wang, Q., Jiang, S., Pan, X., Wang, F., Yang, H., Bolund, L., Lin, C., Han, P. & Luo, Y. (2023). Recent development of urinary biomarkers for bladder cancer diagnosis and monitoring. Clinical and Translational Discovery, 3(2), Article e183. https://doi.org/10.1002/ctd2.183

Zeng, Y., Lv, W., Tao, H., Li, C., Jiang, S., Liang, Y., Chen, C., Yu, T., Li, Y., Wu, S., Cui, X., Liang, N., Wang, P., Xu, H., Dong, J., Teng, H., Chen, K., Mu, K., Fan, T. ... Han, P. (2025). Mapping the chromothripsis landscape in urothelial carcinoma unravels great intratumoral and intertumoral heterogeneity. iScience, 28(1), 111510. Article 111510. https://doi.org/10.1016/j.isci.2024.111510

Yuan, H., Song, Z., Sun, X.-Q., Song, C., Guo, L., Zhang, Q., Luo, Y., Yuan, C., Gao, J. & Jia, X. (2025). Profiling the cell diversity and tissue structure of aqueous humor circulatory system in human eyes using spatial single-cell RNA sequencing. Genes & diseases, 12(1), 101304. Article 101304. https://doi.org/10.1016/j.gendis.2024.101304

Yuan, H., Song, C., Xu, H., Sun, Y., Anthon, C., Bolund, L., Lin, L., Benabdellah, K., Lee, C., Hou, Y., Gorodkin, J. & Luo, Y. (2025). An Overview and Comparative Analysis of CRISPR-SpCas9 gRNA Activity Prediction Tools. The CRISPR Journal, 8(2), 89-104. https://doi.org/10.1089/crispr.2024.0058

Yu, J., Xiang, X., Huang, J., Liang, X., Pan, X., Dong, Z., Petersen, T. S., Qu, K., Yang, L., Zhao, X., Li, S., Zheng, T., Xu, Z., Liu, C., Han, P., Xu, F., Yang, H., Liu, X., Zhang, X. ... Lin, L. (2020). Haplotyping by CRISPR-mediated DNA circularization (CRISPR-hapC) broadens allele-specific gene editing. Nucleic Acids Research, 48(5), Article e25. https://doi.org/10.1093/nar/gkz1233

YeYang, S. U., Lin, L., Geng, TIAN., Chen, CHEN., Tao, LIU., XingYa, XU., XinPeng, QI. & XiuQing, ZHANG. (2009). Prepare re-sequencing DNA library of 2 cancer candidate genes using ligation-by-amplification protocol by two PCR reactions. Science in China, Series C: Life Sciences, 52(5), 483-491. https://doi.org/10.1007/s11427-009-0066-8

Yang, S., Lan, T., Wei, R., Lin, L., Du, H., Huang, Y., Zhang, G., Huang, S., Shi, M., Wang, C., Li, R., Han, L., Tang, D., Li, H., Zhang, H., Cui, J., Lu, H., Huang, J., Luo, Y. ... Fang, S.-G. (2023). Single-nucleus transcriptome inventory of giant panda reveals cellular basis for fitness optimization under low metabolism. BMC Biology, 21(1), 222. Article 222. https://doi.org/10.1186/s12915-023-01691-2

Xu, Z., He, J., Han, P., Dai, P., Lv, W., Liu, N., Liu, L., Liu, L., Pan, X., Xiang, X., Li, H., Ge, F., Gao, S., Liao, Z., Luo, Y. & Li, Y. (2023). Plasma extrachromosomal circular DNA is a pathophysiological hallmark of short-term intensive insulin therapy for type 2 diabetes. Clinical and Translational Medicine, 13(10), Article e1437. https://doi.org/10.1002/ctm2.1437

Xiong, K., Zhou, Y., Hyttel, P., Bolund, L., Freude, K. K. & Luo, Y. (2016). Generation of induced pluripotent stem cells (iPSCs) stably expressing CRISPR-based synergistic activation mediator (SAM). Stem Cell Research, 17(3), 665-669. https://doi.org/10.1016/j.scr.2016.10.011

Xiong, K., Zhou, Y., Blichfeld, K. A., Hyttel, P., Bolund, L., Freude, K. K. & Luo, Y. (2017). RNA-Guided Activation of Pluripotency Genes in Human Fibroblasts. Cellular Reprogramming, 19(3), 189-198. https://doi.org/10.1089/cell.2017.0006

Xiao, J., Wang, F., Hu, X., Li, D., Liu, G., Xu, Q., Chen, C., Xiang, H., Dong, X., Zhu, L., Yang, D., Gao, Y., Wang, M., Luo, Y., Chao, C. C., Li, G. & Guo, Q. (2023). Engineering In Vitro Organ-Structured Tumor Model for Evaluating Neoantigen-Specific T Cell Responses in Hepatocellular Carcinoma. Advanced Materials Interfaces, 10(24), Article 2300155. https://doi.org/10.1002/admi.202300155

Xiang, X., Luo, L., Nodzyński, M., Li, C., Han, P., Dou, H., Petersen, T. S., Liang, X., Pan, X., Qu, K., Yang, L., Dang, Y., Liu, X., Bolund, L., Zhang, X., Tong, G., Xing, Y., Luo, Y. & Lin, L. (2019). LION: a simple and rapid method to achieve CRISPR gene editing. Cellular and Molecular Life Sciences, 76(13), 2633-2645. https://doi.org/10.1007/s00018-019-03064-x

Xiang, X., Li, C., Chen, X., Dou, H., Li, Y., Zhang, X. & Luo, Y. (2019). CRISPR/Cas9-Mediated Gene Tagging: A Step-by-Step Protocol. In Y. Luo (Ed.), Methods in Molecular Biology (pp. 255-269). Humana Press. https://doi.org/10.1007/978-1-4939-9170-9_16

Xiang, X., Zhao, X., Pan, X., Dong, Z., Yu, J., Li, S., Liang, X., Han, P., Qu, K., Jensen, J. B., Farup, J., Wang, F., Petersen, T. S., Bolund, L., Teng, H., Lin, L. & Luo, Y. (2021). Efficient correction of Duchenne muscular dystrophy mutations by SpCas9 and dual gRNAs. Molecular Therapy - Nucleic Acids, 24, 403-415. https://doi.org/10.1016/j.omtn.2021.03.005

Xiang, X., Corsi, G. I., Anthon, C., Qu, K., Pan, X., Liang, X., Han, P., Dong, Z., Liu, L., Zhong, J., Ma, T., Wang, J., Zhang, X., Jiang, H., Xu, F., Liu, X., Xu, X., Wang, J., Yang, H. ... Luo, Y. (2021). Enhancing CRISPR-Cas9 gRNA efficiency prediction by data integration and deep learning. Nature Communications, 12(1), Article 3238. https://doi.org/10.1038/s41467-021-23576-0

Wang, J., Luo, Y., Wang, K., Wang, Y., Zhang, X., Teng, H. & Sun, Z. (2014). Clock-controlled StAR's expression and corticosterone production contribute to the endotoxemia immune response. Chronobiology International, 1-10. https://doi.org/10.3109/07420528.2014.982284

Wang, J., Zhang, X., Cheng, L. & Luo, Y. (2020). An overview and metanalysis of Machine and Deep Learning-Based CRISPR gRNA Design Tools. RNA Biology, 17(1), 13-22. https://doi.org/10.1080/15476286.2019.1669406

Wang, J., Xiang, X., Bolund, L., Zhang, X., Cheng, L. & Luo, Y. (2020). GNL-Scorer: A generalized model for predicting CRISPR on-target activity by machine learning and featurization. Molecular and Cellular Biology, 12(11), 909-911. https://doi.org/10.1093/jmcb/mjz116

Wang, F., Xu, Q., Zhuang, Z., Li, Z., Gao, Q., Huang, Y., Luo, Y., Zhang, X., Zhu, L. & Chao, C.-C. (2021). A single-cell approach to engineer CD8+ T cells targeting cytomegalovirus. Cellular & molecular immunology, 18, 1326-1328. https://doi.org/10.1038/s41423-020-0466-z

Wang, F., Ding, P., Liang, X., Ding, X., Brandt, C. B., Sjöstedt, E., Zhu, J., Bolund, S., Zhang, L., de Rooij, L. P. M. H., Luo, L., Wei, Y., Zhao, W., Lv, Z., Haskó, J., Li, R., Qin, Q., Jia, Y., Wu, W. ... Luo, Y. (2022). Endothelial cell heterogeneity and microglia regulons revealed by a pig cell landscape at single-cell level. Nature Communications, 13(1), Article 3620. https://doi.org/10.1038/s41467-022-31388-z

Wang, F., Ding, P., Liang, X., Ding, X., Brandt, C. B., Sjöstedt, E., Zhu, J., Bolund, S., Zhang, L., de Rooij, L. P. M. H., Luo, L., Wei, Y., Zhao, W., Lv, Z., Haskó, J., Li, R., Qin, Q., Jia, Y., Wu, W. ... Luo, Y. (2022). Author Correction: Endothelial cell heterogeneity and microglia regulons revealed by a pig cell landscape at single-cell level (Nature Communications, (2022), 13, 1, (3620), 10.1038/s41467-022-31388-z). Nature Communications, 13(1), Article 6748. https://doi.org/10.1038/s41467-022-34498-w

Wang, Z., Song, C., Hong, X., Yu, J., Xu, Z., Han, P., He, C., Mao, S., Li, Z., Yuan, H., Wang, X., Zhai, J., Tan, S., Fan, W., Xu, Y., Zhang, T., Yang, Z., Lin, S., Zhu, W. ... Xiang, X. (2026). Reduced eccDNA content in idiopathic asthenozoospermia sperm is associated with compromised DNA repair capacity and elevated DNA damage. Cellular & Molecular Biology Letters, 31(1), Article 57. https://doi.org/10.1186/s11658-026-00857-4

Wang, F., Ren, J., Zhou, Y., Huo, W., Liu, H., Qin, M., Jing, Y., Yin, Z., Shi, T., Shan, H., Tan, G., Wang, H., Guo, X., Mei, J., Yue, Z., Hou, Z., Zhou, H., Song, J., Degn, S. E. ... Jiang, Y. (2026). Cross-species comparison of amniote single-cell transcriptomes reveals evolutionary conservation and divergence in the chicken immune system. Nature Communications, 17(1). https://doi.org/10.1038/s41467-026-72642-y

Vad-Nielsen, J., Lin, L., Bolund, L., Nielsen, A. L. & Luo, Y. (2016). Golden Gate Assembly of CRISPR gRNA expression array for simultaneously targeting multiple genes. Cellular and Molecular Life Sciences, 73(22), 4315–4325. https://doi.org/10.1007/s00018-016-2271-5

Vad-Nielsen, J., Lin, L., Jensen, K. T., Nielsen, A. L. & Luo, Y. (2016). A Golden Gate-based Protocol for Assembly of Multiplexed gRNA Expression Arrays for CRISPR/Cas9. Bio-protocol, 6(23), Article e2059. https://doi.org/10.21769/BioProtoc.2059

Vad-Nielsen, J., Nielsen, A. L. & Luo, Y. (2018). Simple method for assembly of CRISPR synergistic activation mediator gRNA expression array. Journal of Biotechnology, 274, 54-57. https://doi.org/10.1016/j.jbiotec.2018.03.018

Toustrup, L. B., Zhou, Y., Kvistgaard, H., Gregersen, N., Rittig, S., Aagaard, L., Corydon, T. J., Luo, Y. & Christensen, J. H. (2017). Induced pluripotent stem cells derived from a patient with autosomal dominant familial neurohypophyseal diabetes insipidus caused by a variant in the AVP gene. Stem Cell Research, 19, 37-42. https://doi.org/10.1016/j.scr.2016.12.021

Thomsen, E. A., Rovsing, A. B., Anderson, M. V., Due, H., Huang, J., Luo, Y., Dybkaer, K. & Mikkelsen, J. G. (2020). Identification of BLNK and BTK as mediators of rituximab-induced programmed cell death by CRISPR screens in GCB-subtype diffuse large B-cell lymphoma. Molecular Oncology, 14(9), 1978-1997. https://doi.org/10.1002/1878-0261.12753

Teuwen, L. A., De Rooij, L. P. M. H., Cuypers, A., Rohlenova, K., Dumas, S. J., García-Caballero, M., Meta, E., Amersfoort, J., Taverna, F., Becker, L. M., Veiga, N., Cantelmo, A. R., Geldhof, V., Conchinha, N. V., Kalucka, J., Treps, L., Conradi, L. C., Khan, S., Karakach, T. K. ... Carmeliet, P. (2021). Tumor vessel co-option probed by single-cell analysis. Cell Reports, 35(11), Article 109253. https://doi.org/10.1016/j.celrep.2021.109253

The PigGTEx Consortium (2024). A compendium of genetic regulatory effects across pig tissues. Nature Genetics, 56(1), 112-123. https://doi.org/10.1038/s41588-023-01585-7

Sun, Y., Qu, K., Corsi, G. I., Anthon, C., Pan, X., Xiang, X., Jensen, L. J., Lin, L., Luo, Y. & Gorodkin, J. (2025). Deep learning models simultaneously trained on multiple datasets improve base-editing activity prediction. Nature Communications, 16(1), Article 9821. https://doi.org/10.1038/s41467-025-65200-5

Song, Q., Datta, S., Liang, X., Xu, X., Pavicic, P., Zhang, X., Zhao, Y., Liu, S., Zhao, J., Xu, Y., Xu, J., Wu, L., Wu, Z., Zhang, M., Zhao, Z., Lin, C., Wang, Y., Han, P., Jiang, P. ... Yang, J. (2023). Type I interferon signaling facilitates resolution of acute liver injury by priming macrophage polarization. Cellular & molecular immunology, 20(2), 143-157. https://doi.org/10.1038/s41423-022-00966-y

Sølvsten, C., Daugaard, T. F., Luo, Y., de Paoli, F. V., Christensen, J. H. & Nielsen, A. L. (2017). The Effects of Voluntary Physical Exercise-Activated Neurotrophic Signaling in Rat Hippocampus on mRNA Levels of Downstream Signaling Molecules. Journal of Molecular Neuroscience, 62(2), 142-153. https://doi.org/10.1007/s12031-017-0918-9

Skorda, A., Lauridsen, A. R., Wu, C., Huang, J., Mrackova, M., Winther, N. I., Jank, V., Sztupinszki, Z., Strauss, R., Bilgin, M., Maeda, K., Liu, B., Luo, Y., Jäättelä, M. & Kallunki, T. (2023). Activation of invasion by oncogenic reprogramming of cholesterol metabolism via increased NPC1 expression and macropinocytosis. Oncogene, 42(33), 2495-2506. https://doi.org/10.1038/s41388-023-02771-x

Sjöstedt, E., Zhong, W., Fagerberg, L., Karlsson, M., Mitsios, N., Adori, C., Oksvold, P., Edfors, F., Limiszewska, A., Hikmet, F., Huang, J., Du, Y., Lin, L., Dong, Z., Yang, L., Liu, X., Jiang, H., Xu, X., Wang, J. ... Mulder, J. (2020). An atlas of the protein-coding genes in the human, pig, and mouse brain. Science, 367(6482), Article 5947. https://doi.org/10.1126/science.aay5947

Shi, M., Méar, L., Karlsson, M., Álvez, M. B., Digre, A., Schutten, R., Katona, B., Vuu, J., Lindström, E., Hikmet, F., Jin, H., Yuan, M., Li, X., Yang, H., Song, X., Sjöstedt, E., Edfors, F., Oksvold, P., von Feilitzen, K. ... Zhang, C. (2025). A resource for whole-body gene expression map of human tissues based on integration of single cell and bulk transcriptomics. Genome Biology, 26(1), 152. Article 152. https://doi.org/10.1186/s13059-025-03616-4

Secher, J. O., Ceylan, A., Mazzoni, G., Mashayekhi, K., Li, T., Muenthaisong, S., Nielsen, T. T., Li, D., Li, S., Petkov, S., Cirera, S., Luo, Y., Thombs, L., Kadarmideen, H. N., Dinnyes, A., Bolund, L., Roelen, B. A. J., Schmidt, M., Callesen, H. ... Freude, K. K. (2017). Systematic in vitro and in vivo characterization of Leukemia-inhibiting factor- and Fibroblast growth factor-derived porcine induced pluripotent stem cells. Molecular Reproduction and Development, 84(3), 229-245. https://doi.org/10.1002/mrd.22771

Secher, J., Liu, Y., Petkov, S., Luo, Y., Li, D., Hall, V. J., Schmidt, M., Callesen, H., Bentzon, J. F., Sørensen, C. B., Freude, K. K. & Hyttel, P. (2017). Evaluation of porcine stem cell competence for somatic cell nuclear transfer and production of cloned animals. Animal Reproduction Science, 178, 40-49. https://doi.org/10.1016/j.anireprosci.2017.01.007

Schmidt, M., Kragh, P. M., Li, J., Lin, L., Liu, Y., Vajta, G. & Callesen, H. (2010). Establishment of Pregnancies in Large White Sow Recipients after Transfer of Cloned Embryos of Different Pig Breeds. Reproduction, Fertility and Development, (1), 197-198.

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Yonglun Luo